I have two theories: one says Wekono and the other says Seris. I do know whoever it is, they are opposed to the Void based on some of his voicelines. I would suggest Wekono is the more likely of the two, based not just on the color of his crystal, but also because if she has enough of a stake in this fight to summon Mal'Damba to fight for her, it's more than likely she has enough of a stake to lay a curse on Androxus. Which would mean that Wekono should theoretically be fighting for the Resistance instead of being neutral (according to the current lore), but whatever.
Probably just Androxus as is.
In the OG lore he was cursed after killing Nox (Nyx). She is the Roman/Greek goddess of the night. That was back when Paladins lore was more intertwined with Smite's lore, which is how we got Grover in Paladins (being that he's also Sylvanus' treant).